(Outgroup,(Paramesturus,(Mesturus,Micropycnodon,(Gyrodus,(Arduafrons,((Eomesodon_barnesi,Eomesodon_liassicus,((((Stenamara,(Stemmatodus,((Tepexichthys,(Iemanja,((Coelodus_saturnus,((Nursallia_veronae,(Nursallia_gutturosum,(Abdobalistum,(Nursallia_goedeli,Palaeobalistum_orbiculatum)))),(Oropycnodus_ponsorti,Pycnodus))),(Neoproscinetes,Proscinetes)))),(Anomoeodus,Ocloedus_subdiscus)))),(Macromesodon_cf_M_bernissartensis,(Macromesodon_bernissartensis,Macromesodon_macropterus))),(Trewavasia,(Coccodus,Ichthyoceros))),(Apomesodon_gibbosus,Apomesodon_surgens))),(Brembodus,Gibbodon)))))));