(Outgroup,(Preondactylus,((Dimorphodon,Peteinosaurus),((Batrachognathus,(Versperopterus,(Jeholopterus,(Anurognathus,Dendrorhynchoides)))),(((Austriadactylus,Raeticodactylus),(Campylognathoides,Eudimorphodon)),(Fenghuangopterus,((Sordes,(Scaphognathus,(Pterorhynchus,(Dorygnathus,(Nesodactylus,Rhamphorhynchus))))),(Cacibupteryx,(Changchengopterus,(Mythunga,(Darwinopterus,((((Pterodactylus,Cycnorhamphus),(Cathayopterus,(Ctenochasma,(Eosipterus,(Pterodaustro,(Gegepterus,(P_longicolum,Gnathosaurus))))))),((G_rhamphastinus,(G_cristatus,(Noripterus,(Dsungaripterus,Tatal_pterosaur)))),((((Lonchodectes,(Tupuxuara,Thalassodromeus)),(Zhejiangopterus,(Quetzalcoatlus,Azhdarcho))),(Shenzhoupterus,Chaoyangopterus)),(Tupandactylus,(Tapejara,(Sinopterus,Huaxiapterus)))))),(((Ludodactylus,(Cearadactylus,Guidraco)),((((Haopterus,(Istiodactylus,Nurhachius)),(Ferrodraco,(Ornithocheirus,(Anhanguera,Coloborhynchus)))),(Pteranodon,(Nyctosaurus,Muzquizopteryx))),(Boreopterus,Zhenyuanopterus))),(((Moganopterus,Elanodactylus),(Feilongus,Beipiaopterus)),(Gladocephaloideus,Pterofiltrus)))))))))))))));