(Cytheridella_chariessa,((Leucocythere,Limnocythere_dadayi),(Gomphocythere_ortali,(Gomphocythere_parcedilatata,(Gomphocythere_capensis,((Gomphocythere_aethiopis,Gomphocythere_obtusata),((Gomphocythere_downingi,(Gomphocythere_curta,(Gomphocythere_lenis,(Gomphocythere_simplex,(Gomphocythere_angusta,(Gomphocythere_cristata,(Gomphocythere_alata,(Gomphocythere_coheni,Gomphocythere_woutersi)))))))),(Gomphocythere_angulata,Gomphocythere_wilsoni))))))));