(L_diagnosticus,(H_foxii,((Z_robustus,(D_altus,(C_dispar,((I_atherfieldensis,I_bernissartensis),(O_nigeriensis,((P_byrdi,(P_mongoliensis,(B_johnsoni,E_regalis))),(A_kurzanovi,E_caroljonesa))))))),(T_tilletti,T_dossi))));