(Hyracotherium,(Pachynolophus,(Danjiangia,(Lambdotherium,(Eotitanops,(cf_Eotitanops,(Palaeosyops,(Bunobrontops_savagei,(Mesatirhinus_junius,((Dolichorhinus_hyog,(Desmatotitan_tukhum,Acrotitan_ulanshireh)),(Sphenocoelus_uinten,((Microtitan_mongol,(Fossendorhinus_diplo,(Metarhinus_pater,(Metarhinus_fluviatilis,Metarhinus_abbotti)))),((Sthenodectes_incisivum,(Wickia_brevirhinus,Metatelmatherium_ulti)),(Telmatherium_validus,((Qufutitan_zhoui,Nanotitanops_shang),(Epimanteoceras_formosus,((Protitan_grangeri,Protitan_minor),((Protitanotherium_emarg,(Diplacodon_elatus,(Pachytitan_ajax,(Parabrontops_gobiensis,(Parvicornis_occident,(Eubrontotherium_clarn,(Protitanops_curryi,(Dianotitan_lunanensis,(Duchesneodus_uintensis,(Megacerops_kuwagata,(Notiotitanops_mississ,Megacerops_coloradensis))))))))))),((Rhinotitan_kaiseni,Rhinotitan_andrewsi),((Aktautitan_hippopot,(Pollyosbornia_altidens,Gnathotitan_berkeyi)),((Pygmaetitan_panxianen,((Brachydiast_transylv,Metatitan_khaitshinus),(Metatitan_primus,Metatitan_relictus))),(Nasamplus_progressus,(Protembolotherium_efr,(Embolotherium_andrewsi,Embolotherium_grangeri))))))))))))))))))))))));