(Hyracotherium,(Pachynolophus,(Danjiangia,(Lambdotherium,(Eotitanops,(cf_Eotitanops,(Palaeosyops,(Bunobrontops_savagei,(Mesatirhinus_junius,((Dolichorhinus_hyognathus,(Desmatotitan_tukhumensis,Acrotitan_ulanshirehensis)),(Sphenocoelus_uintensis,((Microtitan_mongoliensis,(Fossendorhinus_diploconus,(Metarhinus_fluviatilis,Metarhinus_abbotti))),(Sthenodectes_incisivum,((Telmatherium_validus,(Qufutitan_zhoui,(Wickia_brevirhinus,Metatelmatherium_ultimum))),((Epimanteoceras_formosus,Nanotitanops_shanghuangensis),((Protitan_grangeri,Protitan_minor),(Protitanotherium_emarginatum,((Rhinotitan_kaiseni,Rhinotitan_andrewsi),(((Aktautitan_hippopotamopus,(Pollyosbornia_altidens,Gnathotitan_berkeyi)),((Pygmaetitan_panxianensis,(Nasamplus_progressus,(Protembolotherium_efremovi,(Embolotherium_andrewsi,Embolotherium_grangeri)))),((Brachydiastematherium_transylvanicum,Metatitan_khaitshinus),(Metatitan_primus,Metatitan_relictus)))),((Pachytitan_ajax,Diplacodon_elatus),(Parabrontops_gobiensis,((Eubrontotherium_clarnoensis,Parvicornus_occidentalis),(Protitanops_curryi,(Notiotitanops_mississippiensis,(Duchesneodus_uintensis,(Dianotitan_lunanensis,(Megacerops_coloradensis,Megacerops_kuwagatarhinus)))))))))))))))))))))))))));