(Hyracotherium,(Pachynolophus,(Danjiangia_pingi,(Lambdotherium,(Eotitanops,(cf_Eotitanops_Eberle_2006,(Palaeosyops,(Bunobrontops_savegei,(Mesatirhinus_junius,(Dolichorhinus_hyognathus,((Desmatotitan_tukhumensis,Acrotitan_ulanshirehensis),(Sphenocoelus_uintensis,((Microtitan_mongoliensis,(Fossendorhinus_diploconus,(Metarhinus_abbotti,(Metarhinus_pater,Metarhinus_fluviatilis)))),(Sthenodectes_incisivum,((Telmatherium_validus,(Wickia_brevirhinus,Metatelmatherium_ultimum)),((Qufutitan_zhoui,Nanotitanops_shanghuanensis),(Epimanteoceras_formosus,((Protitan_grangeri,Protitan_minor),(((Rhinotitan_kaiseni,Rhinotitan__andrewsi),(Pygmaetitan_panxianensis,((Aktautitan_hippopotamopus,(Pollyosbornia_altidens,Gnathotitan_berkeyi)),((Metatitan_khaitshinus,(Brachydiastematherium_transylvanicum,(Metatitan_primus,Metatitan_relictus))),(Nasamplus_progressus,(Protembolotherium_efremovi,(Embolotherium_andrewsi,Embolotherium_grangeri))))))),(Protitanotherium_emarginatum_sensu_stricto,(TMM_41723_dash_3,(Sthenodectes_australis,(Protitanotherium_emarginatum_sensu_lato,((Pachytitan_ajax,(Diplacodon_elatus,Diplacodon_gigan)),(Parabrontops_gobiensis,(Parvicornus_occidentalis,(Eubrontotherium_clarnoensis,(Protitanops_curryi,(Duchesneodus_uintensis,(Dianotitan_lunanensis,(Megacerops_kuwakatarhinus,(Notiotitanops_mississippiensis,Megacerops_coloradensis))))))))))))))))))))))))))))))));