(Outgroup,((Holoptychius,(Hadronector,Allenypterus)),(Polypterus,((Cheirolepis,(Howqualepis,(Moythomasia,((Pteroniscus,(Palaeoniscus,(Eserratus,Paramesolepis))),(Cyranorhis,(Wendyicthys,Aesopichthys)))))),(Paratarrasius,Tarrasius)))));