(Outgroup,(Preondactylus,((Dimorphodon,Peteinosaurus),((Austriadactylus,((Raeticodactylus,(Campylognathoides,Eudimorphodon)),(((Scaphognathus,Sordes),(Pterorhynchus,(Cacibupteryx,Dorygnathus,Angustinaripterus,(Sericipterus,(Nesodactylus,Rhamphorhynchus,Qinglongopterus))))),(Darwinopterus,Pterodactylus)))),(Anurognathus,Batrachognathus,(Dendrorhynchoides,Jeholopterus))))));