(Phenacodus,(Ectocion,((((Propachynolophus_maldani,(Propachynolophus_gaudryi,Propachynolophus_levei,Hyracotherium_leporinum)),(Pliolophus_pernix,Pliolophus_vulpiceps)),(Hyracotherium_sandrae,Cymbalophus_cuniculus)),(Hallensia_louisi,(Pachynolophus_hookeri,Cardiolophus)))));