(Boreosomus_piveteaui,(Birgeria_groelandica,(Chondrosteus_acipenseroides,(Peipiaosteus_pani,((Polyodon_spathula,Protopsephurus_liui),(Priscosturion_longipinnis,(Scaphirhynchus_platorynchus,((Acipenser_oxyrinchus,Acipenser_transmontanus,((Acipenser_baerii,(Huso_huso,Acipenser_ruthenus)),(Acipenser_stellatus,(Pseudoscaphirhynchus_kauffmani,Pseudoscaphirhynchus_hermannni)))),(Acipenser_brevirostrum,Acipenser_fulvescens)))))))));