(Euparkeria,(Herrerasaurus,(Preondactylus,((Dimorphodon,Peteinosaurus),(((Anurognathus,Jeholopterus),(Batrachognathus,Dendrorhynchoides)),(((Austriadactylus,Raeticodactylus),(Pterorhynchus,(((Scaphognathus,Sordes),(Allkaruen_holotype_and_referred,((Darwinopterus,Wukongopterus),(((Pterodactylus,Cycnorhamphus),((Pterodaustro,(Ctenochasma,(Eosipterus,Gegepterus))),(Pterodactylus_longicol,Gnathosaurus))),(((Nurhachius,(Istiodactylus,Haopterus)),((Ludodactylus,(Coloborhynchus,(Ornithocheirus,Anhanguera))),(Boreopterus,(Pteranodon,(Nyctosaurus,Muzquizopteryx))))),((Germanodactylus_rhamph,((Germanodactylus_crist,Noripterus),(Dsungaripterus,Tatal_pterosaur))),(Lonchodectes,((Sinopterus,(Huaxiapterus,(Tapejara,Tupandactylus))),(((Shenzhoupterus,Chaoyangopterus),(Quetzalcoatlus,(Zhejiangopterus,Azhdarcho))),(Tupuxuara,Thalassodromeus)))))))))),(Dorygnathus,(Nesodactylus,(Cacibupteryx,Rhamphorhynchus)))))),(Campylognathoides,Eudimorphodon)))))));