(Zalambdalestes,(Leptictis,((Phenacodus,Meniscotherium),(Protolipterna,(Proterotherium,(Trigonostylops,(Henricosbornia,((Pyrotherium,Notostylops),(Simpsonotus,(((Pleurostylodon,((Puelia,(Eomorphippus,(Eurygenium,(Rhynchippus,(Morphippus,((Colpodon,(Ancylocoelus,(Leontinia,Scarrittia))),(Pascualihippus,(Argyrohippus,((Nesodon,Adinotherium),(Hoffstetterius,(Ponanskytherium,Toxodon))))))))))),(Homalodotherium,Asmodeus))),((Thomashuxleya,Periphragnis),(Pampahippus,Ryphodon))),(Colbertia,((Oldfieldthomasia,(Acropithecus,(Campanorco,(Eohyrax,(Pseudhyrax,((Trachytherus,(Plesiotypotherium,Mesotherium)),(Archaeotypotherium,(Archaeohyrax,((Prohegetotherium,Hegetotherium),(Prosotherium,Paedotherium)))))))))),(Ultrapithecus,(Notopithecus,(Federicoanaya,((Archaeophylus,(Plagiarthrus,(Cochilius,Interatherium))),(Protypotherium,Miocochilius)))))))))))))))));