(Hypothetical_ancestor,(Proganochelys_quenstedti,(Palaeochersis_talampayensis,(Proterochersis_robusta,Australochelys_africanus,((Kayentachelys_aprix,Condorchelys_antiqua),(((Meiolania_platyceps,Mongolochelys_efremovi),(Kallokibotion_bajazidi,(Dorsetochelys_delairi,(Plesiochelys_solodurensis,(Portlandemys_mcdowelli,(Xinjiangchelys_latimarginalis,(Hangaiemys_hoburensis,((((Platysternon_megacephalum,((Gopherus_polyphemus,(Chrysemys_picta,Geoclemys_hamiltonii,(Mongolemys_elegans,((Platychelys_oberndorferi,Caribemys_oxfordiensis,Notoemys_laticentralis),((Elseya_dentata,(Chelodina_oblonga,Phrynops_geoffroanus)),(Pelomedusa_subrufa,(Erymnochelys_madagascariensis,Podocnemis_expansa))))))),((Emarginachelys_cretacea,((Baptemys_wyomingensis,Dermatemys_mawii),(Hoplochelys_crassa,(Staurotypus_triporcatus,(Sternotherus_odoratus,Kinosternon_flavescens))))),((Adocus_beatus,(Zangerlia_neimongolensis,Basilemys_variolosa)),(Peltochelys_durlstonensis,Sandownia_harrisi,((Apalone_spinifera,Lissemys_punctata),(Anosteira_ornata,Carettochelys_insculpta))))))),(Protochelydra_zangerli,(Macroclemys_temminckii,Chelydra_serpentina))),(Toxochelys_latiremis,((Caretta_caretta,Chelonia_mydas),(Mesodermochelys_undulatus,Dermochelys_coriacea)))),(Judithemys_sukhanovi,(Dracochelys_bicuspis,(Sinemys_lens,Ordosemys_leios)))))),(Solnhofia_parsonsi,(Thalassemys_moseri,Santanachelys_gaffneyi)))),((Pleurosternon_bullockii,(Glyptops_plicatulus,Dinochelys_whitei)),(Neurankylus_eximius,(Trinitichelys_hiatti,(Plesiobaena_antiqua,(Boremys_pulchra,(Baena_arenosa,Chisternon_undatum))))))))),(Eileanchelys_waldmani,Heckerochelys_romani)))))));