Contributions > Papers
In press
Payne, A. R. D., Mannion, P. D., Lloyd, G. T. and Davis, K. E., in press.
Decoupling speciation and extinction reveals both abiotic and biotic drivers shaped 250 million years of diversity in crocodile-line archosaurs.
Nature Ecology & Evolution, , .
2023
Hellert, S. M., Grossnickle, D. M., Lloyd, G. T., Kammerer, C. F. and Angielczyk, K. D., 2023.
Derived faunivores are the forerunners of major synapsid radiations.
Nature Ecology & Evolution, , .
metatree GitHub
2022
Wisniewski, A. L., Lloyd, G. T. and Slater, G. J., 2022.
Extant species fail to estimate ancestral geographical ranges at older nodes in primate phylogeny.
Proceedings of the Royal Society of London B, B289, 20212535.
Project GitHub
Dryad upload of supplementary materials
2021
Lloyd, G. T. and Slater, G. J., 2021.
A total-group phylogenetic metatree for Cetacea and the importance of fossil data in diversification analyses.
Systematic Biology, 70, 922-939.
Preprint
Project GitHub
Dryad upload of supplementary materials
Wang, M., Lloyd, G. T., Zhang, C. and Zhou, Z.-H., 2021.
The patterns and modes of the evolution of disparity in Mesozoic Birds.
Proceedings of the Royal Society B, 288, 20203105.
Project GitHub
Dryad upload of code and data
2020
Wright, A. M. and Lloyd, G. T., 2020.
Bayesian analyses in phylogenetic palaeontology: interpreting the posterior sample.
Palaeontology, 63, 997-1006.
Project GitHub
Dryad upload of code and data
Guillerme, T., Cooper, N., Brusatte, S. L., Davis, K. E., Jackson, A. L., Gerber, S., Goswami, A., Healy, K., Hopkins, M. J., Jones, M. E. H., Lloyd, G. T., O'Reilly, J. E., Pate, A., Puttick, M. N., Rayfield, E. J., Saupe, E. E., Sherratt, E., Slater, G. J., Weisbecker, V., Thomas, G. H. and Donoghue, P. C. J., 2020.
Disparities in the analysis of morphological disparity.
Biology Letters, 16, 20200199.
2019
Lehmann, O. E. R., Ezcurra, M. D., Butler, R. J. and Lloyd, G. T., 2019.
Biases with the Generalized Euclidean Distance measure in disparity analyses with high levels of missing data.
Palaeontology, 62, 837-849.
Dryad upload of code and data
2018
Lloyd, G. T., 2018.
Journeys through discrete-character morphospace: synthesizing phylogeny, tempo, and disparity.
Palaeontology, 61, 637-645.
Dryad upload of code and data
Tarver, J. E., Taylor, R. S., Puttick, M. N., Lloyd, G. T., Pett, W., Fromm, B., Schirrmeister, B. E., Pisani, D., Peterson, K. J. and Donoghue, P. C. J., 2018.
Well-annotated microRNAomes do not evidence pervasive miRNA loss.
Genome Biology and Evolution, 6, 1457-1470.
Dunne, E. M., Close, R. A., Button, D. J., Brocklehurst, N., Cashmore, D. D., Lloyd, G. T. and Butler, R. J., 2018.
Diversity change during the rise of tetrapods and the impact of the 'Carboniferous Rainforest Collapse'.
Proceedings of the Royal Society B, 285, 20172730.
2017
Button, D. J., Lloyd, G. T., Ezcurra, M. D. and Butler, R. J., 2017.
Mass extinctions drove increased global faunal cosmopolitanism on the supercontinent Pangaea.
Nature Communications, 8, 733.
2016
Lloyd, G. T., Bapst, D. W., Friedman, M. and Davis, K. E., 2016.
Probabilistic divergence time estimation without branch lengths: dating the origins of dinosaurs, avian flight, and crown birds.
Biology Letters, 12, 20160609.
Dryad upload of code and data
Clarke, J., Lloyd, G. T. and Friedman, M., 2016.
Little evidence for enhanced phenotypic evolution in early teleosts relative to their 'living fossil' sister group.
Proceedings of the National Academy of Sciences USA, 113, 11531-11536.
Bapst, D. W., Wright, A. M., Matzke, N. J. and Lloyd, G. T., 2016.
Topology, divergence dates and macroevolutionary inferences vary between different tip-dating approaches applied to fossil theropods (Dinosauria).
Biology Letters, 12, 20160237.
Dryad upload of data and analytical files
Wright, A. M., Lloyd, G. T. and Hillis, D. M., 2016.
Modeling character change heterogeneity in phylogenetic analyses of morphology through the use of priors.
Systematic Biology, 65, 602-611.
Dryad upload of code and data
Wang, M. and Lloyd, G. T., 2016.
Rates of morphological evolution are heterogeneous in Early Cretaceous birds.
Proceedings of the Royal Society B, 283, 20160214.
Dryad upload of code and data
Lloyd, G. T., 2016.
Estimating morphological diversity and tempo with discrete character-taxon matrices: implementation, challenges, progress, and future directions.
Biological Journal of the Linnean Society, 118, 131-151.
Dryad upload of simulation code and tutorial
Benson, R. B. J., Butler, R. J., Alroy, J., Mannion, P. D., Carrano, M. T. and Lloyd, G. T., 2016.
Near-stasis in the long-term diversification of Mesozoic tetrapods.
PLoS Biology, 14, e1002359.
Dryad upload
2015
Close, R. A., Friedman, M., Lloyd, G. T. and Benson, R. B. J., 2015.
Evidence for a mid-Jurassic adaptive radiation in mammals.
Current Biology, 25, 2137-2142.
Dryad upload of code and data
Wang, M., Zheng, X.-T., O'Connor, J. K., Lloyd, G. T., Wang, X.-L., Wang, Y., Zhang, X.-M. and Zhou, Z.-H., 2015.
The oldest record of Ornithuromorpha from the Early Cretaceous of China.
Nature Communications, 6, 6987.
Bell, M. A. and Lloyd, G. T., 2015.
strap: an R package for plotting phylogenies against stratigraphy and assessing their stratigraphic congruence.
Palaeontology, 58, 379-389.
Dryad upload of R tutorial and example data sets
Brusatte, S. L., Butler, R. J., Barrett, P. M., Carrano, M. T., Evans, D. C., Lloyd, G. T., Mannion, P. D., Norell, M. A., Peppe, D. J., Upchurch, P. and Williamson, T. E., 2015.
The extinction of the dinosaurs.
Biological Reviews, 90, 628-642.
2014
Brusatte, S. L., Lloyd, G. T., Wang, S. C. and Norell, M. A., 2014.
Gradual assembly of avian body plan culminated in rapid rates of evolution across dinosaur-bird transition.
Current Biology, 24, 2386-2392.
Dryad upload of R code and data
2013
Lloyd, G. T. and Friedman, M., 2013.
A survey of palaeontological sampling biases in fishes based on the Phanerozoic record of Great Britain.
Palaeogeography, Palaeoclimatology, Palaeoecology, 372, 5-17.
Dryad upload of R code and data
2012
Lloyd, G. T., Pearson, P. N., Young, J. R. and Smith, A. B., 2012.
Sampling bias and the fossil record of planktonic foraminifera on land and in the deep sea.
Paleobiology, 38, 569-584.
Smith, A. B., Lloyd, G. T. and McGowan, A. J., 2012.
Phanerozoic marine diversity: rock record modelling provides an independent test of large-scale trends.
Proceedings of the Royal Society B, 279, 4489-4495.
Lloyd, G. T., Young, J. R. and Smith, A. B., 2012.
Comparative quality and fidelity of deep-sea and land-based nannofossil records.
Geology, 40, 155-158.
Lloyd, G. T., Wang, S. C. and Brusatte, S. L., 2012.
Identifying heterogeneity in rates of morphological evolution: discrete character change in the evolution of lungfish (Sarcopterygii; Dipnoi).
Evolution, 66, 330-348.
Dryad upload of R code and data
Lloyd, G. T., 2012.
A refined modelling approach to assess the influence of sampling on palaeobiodiversity curves: new support for declining Cretaceous dinosaur richness.
Biology Letters, 8, 123-126.
R function and tutorial
Dryad upload of functions and data
Piece in Adelaide Advertiser describing me as a "leading palaeontologist"
Lloyd, G. T., Young, J. R. and Smith, A. B., 2012.
Taxonomic structure of the fossil record is shaped by sampling bias.
Systematic Biology, 61, 80-89.
Dryad upload of data
2011
Brusatte, S. L., Benton, M. J., Lloyd, G. T., Ruta, M. and Wang, S. C., 2011.
Macroevolutionary patterns in the evolutionary radiation of archosaurs (Tetrapoda: Diapsida).
Transactions of the Royal Society of Edinburgh: Earth and Environmental Science, 101, 367-382.
2010
Sakamoto, M., Lloyd, G. T. and Benton, M. J., 2010.
Phylogenetically structured variance in felid bite force: the role of phylogeny in the evolution of biting performance.
Journal of Evolutionary Biology, 4, 733-736.
2008
Brusatte, S. L., Benton, M. J., Ruta, M. and Lloyd, G. T., 2008.
The first 50 million years of dinosaur evolution: macroevolutionary pattern and morphological disparity.
Biology Letters, 4, 733-736.
Lloyd, G. T., Davis, K. E., Pisani, D., Tarver, J. E., Ruta, M., Sakamoto, M., Hone, D. W. E., Jennings, R. & Benton, M. J., 2008.
Dinosaurs and the Cretaceous Terrestrial Revolution.
Proceedings of the Royal Society B, 275, 2483-2490.
Tree and XML files (zip; 1Mb; published version)
Tree and XML files (zip; 1Mb; 2011 version with correction of minor errors - recommended)
Reduced Baum and Ragan MRP matrix (NEXUS)
Reduced Purvis MRP matrix (NEXUS)
Subsampling code
Research group supertree page
Brusatte, S. L., Benton, M. J., Ruta, M. and Lloyd, G. T., 2008.
Superiority, competition, and opportunism in the evolutionary radiation of dinosaurs.
Science, 321, 1485-1488.
2007
Ruta, M., Pisani, D., Lloyd, G. T. & Benton, M. J., 2007.
A supertree of Temnospondyli: cladogenetic patterns in the most species-rich group of early tetrapods.
Proceedings of the Royal Society B, 274, 3087-3095.
